Rosalind faces a troubling domestic situation. Meanwhile Joyce supervises the antenatal clinic at the hospital, where she meets a worried single mother diagnosed with placenta previa and concerned about raising her child alone.

Rosalind delivers a baby with a suspicious spinal lesion and Dr. Turner suspects spina bifida. Meanwhile, during her district rounds, Joyce is about to visit a difficult patient when a sudden gas explosion ensues chaos and leaves many injured.

A complicated pregnancy leads the Nonnatus team on a path of discovery. Sister Hilda and Dr. Turner get involved with a young woman whose health presents a series of challenges.

Violet must balance her professional and personal views to tackle the issue of meths drinkers. Nurse Crane treats a promising young singer who must decide between two very different futures. Trixie and Matthew’s friendship blossoms into something deeper.

Feel the love as the cooks prepare their favorite comfort food and a dish inspired by a loved one. The cooks pour their hearts and souls into recipes ranging from bacon-wrapped meatloaf and cornbread to crab cakes, elk stew and chicken curry.

Violet and Matthew are worried about Fred’s ability to serve as the mayor’s consort. Later, Fred falls ill after a minor injury. Meanwhile, the Nonnatus team is thrown into turmoil when a newborn baby is diagnosed with bilateral hip dysplasia.

Busy and exhausted, James and Helen can’t get any time together. James thinks he’s found a solution when he hires a new trainee vet, Richard Carmody, but things don’t quite work out as he expects. Mrs. Pumphrey also has a new canine member of her household who has an antisocial problem that she needs addressed urgently. Mrs. Hall takes a big step forward in her relationship with Gerald.

In episode three, some of the South’s most compelling and influential contemporary creators take us to the places that feed their imaginations: Mississippi author Jesmyn Ward, Georgia screenwriter and series creator Michael Waldron, Arkansas songwriter Justin Moore, Mississippi poet Natasha Trethewey, New Orleans songwriter Tarriona “Tank” Ball and Virginia songwriter Thao Nguyen.

James is put to the test with an ailing racehorse. Tristan faces a familiar temptation. Siegfried angles for a prestigious client.

Medical advice clashes with farming wisdom as James and Helen have their first quarrel. Mrs. Pumphrey’s dog gets Tristan in trouble.

Siegfried’s First World War experiences resurface as he struggles to save a racehorse. Helen faces a decision over James’s TB testing plan.

As she sets about helping a mother whose newborn baby is suffering from a clotting disorder, Nancy is harboring a secret of her own. Sister Veronica meets a newlywed on her rounds, and the case shocks the women of Nonnatus house.

In the wake of an uncomfortable encounter, Charlotte does all she can to avoid Sidney, but an emergency unites them. An angry Young Stringer confronts Tom Parker over an injustice. Meanwhile, Tom finds a doctor for the town, but Lady Denham proves hard to win over.

Charlotte and Alexander Colbourne reach an understanding, as she and Colonel Lennox enjoy a close encounter.

Georgiana gets a surprise visit. Colbourne’s shooting party goes ahead and Charlotte has to confront all that she’s lost. Mary and Tom cross swords over his plans for the old town.